The goals of the biology major are to: (1) teach the fundamental
concepts and methods of the biological sciences; (2) engage all students
in science as it is practiced by developing their capacities to use
and evaluate scientific knowledge; and (3) explore relationships
of biology to other sciences and to problems of social, cultural,
and intellectual concern.
By proper selection of biology and other science courses in consultation
with a departmental advisor, the major in biology can prepare a student
for graduate study and research in a wide range of areas including
anatomy, biochemistry, bioengineering, biophysics, botany, cell biology,
conservation biology, developmental biology, ecology, environmental
sciences, immunology, marine biology, microbiology, molecular biology,
mycology, physiology, virology, zoology, etc. The major in biology
can also lead to a variety of careers in applied biology (medicine
and agriculture and their allied fields) and science teaching.
Advanced Placement. Students earning scores of 4 or 5 on the Advanced
Placement Biology exam (AP Bio) will receive four hours of natural
science credit applied toward graduation requirements. Four hours
of biology major credit, designated as Biology 605, are automatically
granted to students earning a score of 5 on the Advanced Placement
exam, or at least 6 on the International Baccalaureate exam. Students
granted AP credit in biology who intend to major in biology are exempted
from taking the introductory sequence (Biology 118, 119, 120) but
must still complete the 27-hour minimum for the major, by earning
23 biology credits in addition to the four granted as Biology 605.
Students with Biology 605 may elect, if they so choose, to take either
Biology 118 plus 119, or Biology 120 for four credits; if both of
these courses are taken, Biology 605 credit will not be granted.
Students with AP credit must still take at least five laboratory
courses.
Exemptions. Exemption (not Placement) tests, offered annually every
August, may be taken by any student wishing to be exempted from taking
Biology 118, 119 and/or 120. Note that a score of 5 in Advanced Placement
Biology (AP Bio) will also exempt any student from taking these courses.
It must be pointed out, however, that these courses are appropriate
and helpful for those intending to take the MCAT or similar proficiency
exams. It is thus recommended that students who foresee taking such
exams take (or, minimally, audit) these courses.
Entry-Level Course Sequence Suggestions. Students with a strong high-school
science background find taking Introductory Biology and Chemistry
courses simultaneously in the first semester both useful and challenging.
Other students benefit greatly from taking only one of these courses
during their first semester at Oberlin. Prospective Biology majors
are urged to consult with a Biology faculty member during Freshman
Advising Period to discuss these options.
Major. The biology major consists of a minimum of 27 hours of course
work in categories I through IV and additional cognate course work
in category V. A minimum of nine hours of advanced course work (defined
as courses in categories II, III and IV) must be completed at Oberlin.
No more than seven hours of the advanced course work may be transfer
credits.
Requirements for the Biology Major
I. Core Biology Courses:
A. Biology 118 (Organismal Biology, lecture) and Biology 119 (Organismal Biology,
lab)
B. Biology 120 (Genetics, Evolution and Ecology, lecture, and lab)
C. Biology 213 (Cell and Molecular Biology, lecture)
II. Upper-level Biology Courses Taken at Oberlin College:
A. At least two 200- or 300-level Biology lecture courses (exclusive of Biology
213)
B. At least one 200- or 300-level Biology lab experience (exclusive of Biology
214)
III. Additional Laboratory Courses:
Two laboratory experiences, one of which may be replaced by a semester of research
taken for credit. (Each lab course or course that includes a lab counts as a
single lab course regardless of the number of lab meetings per week.)
IV. Additional Upper-Level Courses to bring the total number of hours to at least
27:
Requirements in categories III and IV above may be fulfilled by taking 200-
or 300-level courses in Biology; 200- or 300-level courses in Neuroscience; Environmental
Studies 316 (Systems Ecology) and 340 Environmental Systems Modeling); Chemistry
374 (Biochemistry); and Geology 320 (Paleontology). Up to three credit hours
of a 400-level Biology or Neuroscience seminar course and up to three credit
hours of research may be counted toward the major. Honors candidates completing
two full semesters of research may count up to four credit hours toward the major.
Private readings do not count for major credit.
V. Cognate Course Requirements:
A. Chemistry 101 and 102 (or 103 may replace 101 and 102)
B. Chemistry 205
C. Four additional hours selected from the following: Calculus (Math 131, 132
or 133), Chemistry (Chem 254), Physics (103), Statistics (Math 113 or 114).
Minor. There is no minor offered in Biology.
Honors. Candidates for Honors in biology enroll as seniors in BIOL 501 and 502,
carry out a research project in consultation with a faculty committee, write
and defend a thesis based on completed work, and give a required research seminar.
Students interested in this program should initiate a discussion of research
possibilities with any member of the staff during the junior year. Students are
not accepted formally into the Honors Program until their proposed research is
reviewed by the department at the beginning of their first semester of research.
Off-Campus or Summer Study. The department offers limited scholarships for summer
course work at approved field (inland and marine) stations in the United States.
Applications for these scholarships are invited early in the Spring Semester;
awards are made in May. In general, students planning off-campus study are advised
to consult first with their advisor, and then the Chairman to determine acceptability
of courses to the major.
Transfer of Credit. Credits in biology earned elsewhere may be applied to the
Oberlin biology major with the approval of the department chair, as long as 14
of the 27 hour minimum in biology, including nine hours of advanced course work
(defined above, under 'major'), are completed at Oberlin.
Winter Term. Among the topics for Winter Term projects that members of the Biology
Department are willing to sponsor are the following: Mr. Allen: projects relying
upon genetic, molecular biological, and biophysical techniques to address cell
physiological questions. Readings in the physiology of muscle and motile cells.
Instruction and readings in Old Icelandic and Old English literature. Ms. Bennett:
intensive science review; pre-medical internship. Mr. Benzing: the technology
of winemaking and viticulture; tropical biology. Mr. Braford: laboratory research
in comparative neuroanatomy for students who have had Biology 320; neurobiology;
evolution; systematics. Ms. Cruz: projects involving comparative embryology,
developmental biology, marsupial biology. Readings in the history of biology,
philosophy of science, evolution. Ms. Garvin: projects and readings in parasitology,
entomology, ornithology, epidemiology, and ecology. Ms. Laskowski: projects involving
plant physiology, plant molecular biology. Mr. Laushman: projects involving conservation
biology, population genetics, and aquatic vascular plants. Mr. Luck: laboratory
research involving recombinant DNA technology for students who have had Biology
309. Ms. McCormick: projects on the evolution of vertebrate auditory and lateral
line systems; topics on the evolution of vertebrate structure. Mr. Miller: holistic
health; new age/transformational subjects; psychic healing group project; new
paradigms in science; noetic sciences. Mr. Tarvin: projects and readings in vertebrate
evolutionary and behavioral ecology, ornithology, avian blood parasites, molecular
genetic tools for evolutionary analyses and readings in human behavioral ecology
and evolutionary biology. Ms. Thornton: projects involving neuroendocrinology
and behavioral neuroendocrinology. Ms. Treichel: projects involving immunology,
virology, or cancer; health career internships.
Preparation for Post-Graduate Education Medical School, Dental and Veterinary Medicine. Most medical, dental, and veterinary
schools require:
1. One year of General Biology with laboratory.
2. Two years of Chemistry with laboratory.
3. One year of Mathematics.
4. One year of Physics with laboratory.
5. One year of English.
Students planning careers in medicine should consult the Health Career Advisors,
the Office of Career Services, and references in the Science Library for details.
Graduate Schools. Most graduate schools in biology expect a major in the biological
sciences to include:
1. Two years of Chemistry with laboratory.
2. One year of Physics with laboratory.
3. College-level Mathematics (Calculus and Statistics).
4. Courses in Computer Science.
Certain deficiencies in course work may be made up in graduate school but generally
not for graduate credit. Students planning to pursue an advanced degree should
consult individual admission requirements of the schools and graduate departments
in which they are interested. The qualifying examination for most graduate schools
is the Graduate Record Examination (GRE) which consists of two parts, the General
Examination and an Advanced component in biology. The majority of graduate schools
require the General Examination for admission but there is variation with respect
to the Advanced requirement. Students are strongly urged to consult with members
of the Biology Department in preparing for graduate work.
FYSP 116. Field-Based Writing: Ecology of the Vermilion River Watershed
4 hours
2NS, 2HU, WRi Next offered 2005-2006.
FYSP 121. Everyday Evolution
3 hours
3NS, WR
First Semester. For description, please see "First-Year Seminar Program" in
this catalog. Enrollment Limit: 14.
Mr. Laushman
FYSP 156. Biological Advances and Ethical Dilemmas
3 hours
1.5NS, 1.5HU, WR Next offered 2005-2006.
FYSP 165. Feeding the World
3 hours
3NS, QPh, WR Next offered 2005-2006.
FYSP 181. Science From a Bird's Eye View: Ecology, Evolution,
and the Study of Birds
3 hours
3NS, WRi Next offered 2005-2006.
FYSP 182. Traditions of Health and Disease in Folk and
Conventional Medicine
3 hours
3NS, WRi
Second Semester. For description, please see "First-Year Seminar Program" in
this catalog. Enrollment Limit: 14.
Mr. Allen
Introductory
Courses for Non-majors and Prospective Majors
032. Body in Health and Disease
1-2 hours
1-2 NS
First Semester.
Intended primarily for non-science majors, this colloquium explores
how the body functions. Focus of study is shaped by students' interests
and previously has concerned reproductive and sexual health, as well as approaches,
both conventional and alternative, to preventing and treating disease. Enrollment
Limit: 8. Consent of instructor required.
Mr. Allen
115. Field Botany
2 hours
2NS
Second Semester. Second Module. A study of vegetation emphasizing vascular plants
as they occur in Northern Ohio. Identification of species and types of plant
communities and selected aspects of natural history will be emphasized. Prerequisite:
BIOL 118, 119, or consent of instructor. Enrollment Limit: 17.
Mr. Benzing
118. Organismal Biology (Lecture only)
3 hours3NS
First Semester. This course provides prospective biology majors and others with
an integrated introduction to the biology of organisms, from the subcellular
level, through the cellular, tissue, organ, and whole organismal level.
Ms. Cruz, Staff
119. Organismal Biology (Laboratory only)
1 hour
1NS
First Semester. Laboratory exercises will emphasize anatomical and physiological
studies of higher vertebrate and flowering plant whole organisms. Preserved animals
are dissected in some laboratories. Some exercises and discussions are designed
to develop critical thinking and problem-solving skills in data analysis and
interpretation. Enrollment Limit: 24 per section. Note: Students must be concurrently
enrolled in or have already had BIOL 118. Priority will be given to first- and
second-year students.
Ms. Bennett, Ms. Cruz, Staff
120. Genetics, Evolution, and Ecology
4 hours
4NS
Second Semester. This course provides prospective biology majors and others with
an integrated introduction to key biological principles of genetics, ecology,
and evolution. The labs feature exercises and discussions designed to develop
critical thinking and problem-solving skills in data analysis and interpretation.
Note: Priority will be given to first- and second-year students.
Mr. Laushman, Ms. Bennett, Staff
201. Invertebrate Biology
4 hours
4NS
First Semester.
Invertebrates represent over 97 percent of the animal kingdom.
This course is a study of the anatomy, physiology, behavior, and ecology of representatives
of the major phyla and their evolutionary relationships. Lectures will emphasize
functional adaptations of the major organ systems. Laboratories will include
observation, collection, identification and dissection of terrestrial, aquatic,
and marine invertebrates. Prerequisite: BIOL 120. Enrollment Limit: 15.
Staff
203. Vertebrate Structure and Evolution
6 hours
6NS
Second Semester. A study of vertebrate structure from evolutionary and functional
perspectives. This topic is introduced by a discussion of the origin of vertebrates
and their phylogeny and basic embryology. The morphological differences in each
major organ system are studied primarily in terms of phylogenetic history and
functional adaptation. All students must participate fully in the anatomical
laboratory exercises. Prerequisite: BIOL 119 or consent of instructor. Note:
BIOL 120 recommended. Enrollment Limit: 20.
Ms. McCormick
204. Plant Biology
4 hours
4NS
First Semester. An introduction to the anatomy, development, and physiology of
plants with emphasis on flowering plants. Lectures and laboratories will highlight
fundamentals of plant structure and function including adaptive features for
life in specific types of environments. Prerequisite: BIOL 118, 119, 120, or
consent of instructor. Enrollment Limit: 16.
Mr. Benzing, Mr. Miller
205. Community Ecology
4 hours
4NS, WRi Next offered 2005-2006.
206. Population Ecology
4 hours
4NS, QPh, WR
First Semester. Population ecology deals with the biotic and abiotic factors
that influence population size and stability. Variation in these factors leads
to temporal and spatial differences among populations. Major topics include:
population growth; demographic variation and stability; population cycles; metapopulation
dynamics; and management considerations. Labs will include field trips to study
natural populations, and computer modeling to understand population dynamics.
Prerequisite: BIOL 120 and consent of instructor. Enrollment Limit: 14.
Mr. Laushman
207. The Fungi
3 hours
3NS
First Semester. First Module. A study of the major groups of the fungi. Lectures
and laboratory will deal with the ecological and economic importance, the life
cycles, and evolutionary relationships of the fungi. Laboratories will emphasize
the collection and identification of local edible and poisonous species of mushrooms
with weekly Tuesday field trips. Thursday labs will be devoted to identifying
new specimens and examination of important, smaller, less well-known fungi. Prerequisites:
BIOL 118, 119, and 120 or consent of instructor. Enrollment Limit: 16.
Mr. Miller
211. Genetics
3 hours
3NS Next offered 2005-2006.
212. Behavioral Ecology
4 hours
4NS
Second Semester. Behavioral ecology is the study of how behavior (broadly defined)
influences lifetime reproductive success of individuals. It is therefore deeply
rooted in evolutionary theory, and considers the ways in which behavior may be
adaptive. Topics will include life history strategies, optimal foraging and habitat
selection, signaling and communication, sexual selection and mating strategies,
and social organization and cooperation. Lectures, laboratories, and discussion
sessions will primarily emphasize field studies and will illustrate observational,
experimental, comparative, and modeling approaches. Laboratories occasionally
meet at night. Prerequisite: BIOL 120. Enrollment Limit: 24. Note: BIOL 212 meets
during alternate spring semesters.
Staff
213. Cell and Molecular Biology (Lecture only)
3 hours
3NS
First Semester. A detailed study of cell structure and function at the molecular
level. Topics to be discussed include structure and function of membranes and
cell organelles; gene structure, function and regulation; bioenergetics; cell
cycle control, signal transduction and genetic engineering. The intent of the
course is to integrate molecular biology, biochemistry, and cell biology in order
to provide a firm foundation for many of the more specialized courses in the
major. Prerequisites: BIOL 118, 119, or 120, and CHEM 102 or 103. Note: Not recommended
for first-year students.
Mr. Luck, Ms. Treichel
214. Cell and Molecular Biology (Laboratory only)
1 hour
1NS
First Semester. Laboratory exercises are designed to illustrate processes central
to cell and molecular biology and to familiarize students with basic skills required
at the laboratory bench. Co-requisite: BIOL 213. Enrollment Limit: 20 per section.
Mr. Luck, Ms. Thornton, Ms. Treichel, Ms. Romberg
215. Ornithology
3 hours
3NS Next offered 2005-2006.
216. Ornithology Laboratory
1 hour
1NS Next offered 2005-2006.
218. Evolution
3 hours
3NS
First Semester. Principles of microevolution (selection, gene flow, mutation,
genetic drift, and factors that influence population genetic structure), evolutionary
ecology (life history strategies, gene-environment interactions), and macroevolution
(changes above the species level) will be studied, with emphasis on both the
process and pattern of organic evolution. Prerequisites: BIOL 118, 120. Consent
of instructor required. Enrollment Limit: 24.
Staff
301. Developmental Biology
3 hours
3NS, WR
Second Semester. A survey of the developmental processes at the molecular, cellular
and organismal levels in vertebrate and invertebrate animals, and also certain
plants and protists. Two 50-minute lectures will explore cell differentiation,
pattern formation, and morphogenesis in embryos, presented with an emphasis on
evolution. A three-hour demonstration-discussion session per week will complement
lecture material with specimens, models and student-led presentations. Prerequisites:
BIOL 118, 119 and 213; or consent of instructor. Enrollment
Limit: 15.
Ms. Cruz
302. Developmental Biology Laboratory
2 hours
2NS
Second Semester. Experiments and laboratory exercises designed to familiarize
students with approaches to the study of embryogenic and developmental processes
at the cell, tissue, and organismal levels. Live and preserved animals will be
used in class, which will meet for one-half to two hours per week in addition
to regularly scheduled class times. Pre- or co-requisite: BIOL 301. Enrollment
Limit: 15.
Ms. Cruz
304. Mechanisms of Plant Adaptation
3 hours
3NS
Second Semester. This course focuses on cellular and molecular mechanisms that
affect plants' ability to succeed in natural and agricultural populations.
Topics will include central issues of plant physiology such as control of flowering,
and nitrogen use, which have implications for agriculture, as well other issues
of ecological significance such as natural defenses against plant pathogens,
response to stresses such as cold and salinity, and mechanisms of light perception.
Creation of transgenic plants for use in agriculture and research will be discussed.
Prerequisite: BIOL 213 or consent of the instructor.
Ms. Laskowski
305. Experiments in Plant Growth and Development
2 hours
2NS
Second Semester. In this laboratory course, students will learn some of the molecular
and genetic techniques currently used in plant research and employ them in independent
projects. Working together, we will choose a set of novel projects that can be
carried out over the course of the semester. Because we will work with living
organisms, and use experimental techniques that do not always fit into three-hour
labs, students will be required to work independently for three to four hours
per week outside of scheduled lab times to complete their projects. Co-requisite:
BIOL 304. Enrollment Limit: 8.
Ms. Laskowski
306. Microbiology (Lecture only)
3 hours
3NS
Second Semester.
The study of microorganisms with an emphasis on prokaryotes.
Major areas to be covered include microbial cell structure and function, physiology,
metabolism, genetics, diversity, and ecology. Applied aspects of microbiology
will also be discussed such as biotechnology, the role of microorganisms in environmental
processes, and medical microbiology. Prerequisites: BIOL 214 and CHEM 205. Enrollment
Limit: 25.
Ms. Romberg
307. Microbiology (Laboratory)
1 hour
1NS
Second Semester. Laboratory exercises are designed to illustrate processes central
to microbiology and to familiarize students with basic skills required for working
with microorganisms. Topics will include asceptic technique, microscopy, and
isolation and identification of unknown bacteria. Co-requisite. BIOL 306. Enrollment
Limit: 16.
Ms. Romberg
308. Genes and Genetic Engineering (Lecture only)
3 hours
3NS
Second Semester. A lecture course on the chemical, physical and biological properties
of nucleic acids, with emphasis on current topics related to the organization,
replication, transcription, translation, regulation, and manipulation of genetic
material. Prerequisites: BIOL 213 and CHEM 205.
Mr. Luck
309. Genes and Genetic Engineering (Lecture and Laboratory)
4 hours
4NS
Second Semester. Identical to BIOL 308 but with the addition of one laboratory
per week. Laboratories will emphasize techniques used in working with nucleic
acids. Prerequisites: BIOL 214 and CHEM 205. Enrollment Limit: 16.
Mr. Luck
312. Animal Physiology
4 hours
4NS, WR
First Semester. This course explores the function of the body, from the molecular
level (e.g., generation of electrical signals in the nervous system) to the organismal
level (e.g., adaptations to pregnancy, exercise, or extreme environments). Classes
and laboratories study the physiology of excitable cells (e.g., nerves and muscles),
cardiovascular system, lungs and respiratory system, kidneys and renal system,
and reproduction. Prerequisite: BIOL 213 or NSCI 201 or NSCI 204 or consent of
instructor. Enrollment Limit: 28.
Mr. Allen
313. The Living Cell
3 hours
3NS, WR
Second Semester. Course explores the vibrant activity of cells, ranging from
intracellular shuttling of vesicles and organelles to locomotion of cells during
events such as embryogenesis, cancer, or wound healing. To understand how this
activity arises, the course studies the design and function of the involved proteins,
as well as the signaling pathways orchestrating the activity. Involves each week
two class meetings and one small-group discussion. Prerequisite: BIOL 213 or
NSCI 201 or NSCI 204 or consent of instructor. Enrollment Limit: 16.
Mr. Allen
314. The Living Cell (with Research Project)
3-4 hours
3-4NS
Second Semester. Shares two weekly classes with BIOL 313, but includes a semester-long,
small-group research project in place of the small-group discussions. Chosen
by students in consultation with instructor, the projects will develop students' mastery
of current cell biological research techniques (e.g., mutagenesis, genetic screens,
spectroscopy, or advanced microscopy), as well as skills at designing, performing,
and then communicating experiments. Consent of instructor required. Enrollment
Limit: 10.
Mr. Allen
325. Principles of Vascular Plant Taxonomy and Systematics
2 hours
2NS, WR
First Semester. First Module. This course constitutes an introduction to plant
taxonomy and systematics. Lectures, discussions, and readings will focus on the
evolutionary history of vascular plants, mechanisms of speciation, methods for
inferring plant phylogeny and taxonomic nomenclature. The laboratories will be
devoted primarily to familiarization with the information necessary to identify
plants in the field and from preserved specimens. Prerequisite: BIOL 119, 120
or consent of instructor. Enrollment Limit: 24.
Mr. Benzing
326. Vascular Plant Diversity and Systematics
2 hours
2NS, WR
Second Semester. Second Module. This course deals with the diversity of vascular
plants broadly defined. Specific subjects include the adaptive strategies of
higher plants, relationships among the major taxa, particularly families within
the angiosperm complex, and plants as actors within biological communities and
as partners in symbioses. Most of the laboratories will be devoted to field work
and familiarization with the local flora in winter condition and early during
the growing season. Enrollment Limit: 24. Prerequisite: BIOL 119, 120 or consent
of instructor. BIOL 325 highly recommended.
Mr. Benzing
327. Immunology
3 hours
3NS Next offered 2005-2006.
328. Immunology Laboratory
1.5 hours
1.5NS Next offered 2005-2006.
329. Virology
3 hours
3NS
Second Semester. An introduction to the basic principles of virology, including
the biochemistry, molecular genetics, and genetics of viruses with emphasis on
animal viruses. Lectures will examine viruses as important model systems for
elucidating the basic principles of molecular biology and also as important agents
of disease. Medical topics will include a discussion of the pathogenesis, immunology,
and prevention/treatment of important human viral diseases. Prerequisite: BIOL
213. Enrollment Limit: 24.
Ms. Treichel
403. Seminar: Prokaryotic Cell Biology
3 hours
3NS
First Semester. Recent research in microbiology has indicated that bacteria have
far more complicated life cycles and structural organization than had previously
been realized. Principles for achieving this organization are similar to those
used by eukaryotic cells, but the detailed mechanisms often differ substantially.
Topics covered in this course may include bacterial cell structure, the cell
cycle, developmental pathways, response to the enviromnent, motility, and membrane
physiology. Methods for studying prokaryotic cell biology and biochemistry will
be explored through readings from the primary literature. Lectures will alternate
with in-class discussions. Prerequisite: BIOL 213/214. Consent of instructor
required. Preference to juniors and seniors. Enrollment Limit: 12.
Ms. Romberg
501, 502. Research
1-3 hours
1-3NS Projects for original investigation are developed by students in consultation
with a faculty member. Students in the Honors Program enroll for both semesters
of their senior year. A maximum of three credit hours (four hours for Honors
students completing two semesters of research) and one laboratory unit may be
earned in this course towards the requirements for a biology major. Research
sponsored by Mr. Allen, Mr. Benzing, Mr. Braford, Ms. Cruz, Ms. Laskowski, Mr.
Laushman, Mr. Luck, Ms. McCormick, Mr. Miller, Ms. Romberg, Ms. Thornton, and
Ms. Treichel. Consent of instructor required.
995. Private Reading
1-3 hours
1-3NS Independent study of a subject arranged by student with member of Biology teaching
staff, who supervises the project. Only subjects beyond the range of catalog
course offerings permitted. Special approvals required from: project supervisor,
student's academic advisor, and department chairman. Note: A student is
limited to one private reading course per semester. Private readings sponsored
by Mr. Allen, Ms. Bennett, Mr. Benzing, Mr. Braford, Ms. Cruz, Ms. Laskowski,
Mr. Laushman, Mr. Luck, Ms. McCormick, Mr. Miller, Ms. Romberg, Ms. Thornton,
and Ms. Treichel. Consent of instructor required.